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[Glider]

Sounds to me like he had a Recreational licence but was buying and selling a number of animals that was considered excessive by the EPA.

If EPA considered it excessive they would see it as commercial, which he was not licenced for, and the list of commercially tradeable species is much more restricted than those available under a recreational licence, so he would have probably been considered by epa to be commercially trading animals that were not permitted to be commercially traded even if he did have a commercial licence.